and is considered a complete strand. Second, the other strand has a short
RNA at the 5’ end, making it an incomplete strand. Hence, one strand
being complete and the other being incomplete, they have a gapped DNA
genome. The mechanism followed by the members of this group must
consider the facts that the 1) transcription occurs in the host nucleus, 2)
only the RNA part would be exported out of the nucleus, and 3) viral
assembly occurs in the host cytoplasm.
Therefore, the gapped genome is first repaired/filled by viral polymerase forming a
circular DNA. This DNA is transcribed into RNA by host polymerase. The RNA
goes out of the nucleus. Finally, the viral RT enzyme makes DNA from RNA,
making it possible to be packaged into virion capsid.
Example: Hepatitis B (Figure 2.3).
2.3
VIRAL INFECTION CYCLE
The viral genome, despite its limited size, will have to follow the general rule of the
“Central Dogma” to transfer the information from its genes to the proteins. The viral
genome needs to first replicate itself. Second, it must transcribe to form a functional
mRNA, and third, this mRNA must undergo translation to generate a functional
protein, including all post-translational modifications, producing an infectious virion,
and for all these it depends on the host system. An infection cycle needs to follow a
series of events ultimately leading to the production of new virions using the host
cellular machinery. There are three types of commonly known infection cycles.
2.3.1
LYTIC CYCLE
A lytic cycle is also known as virulent infection. This type of infection cycle is mostly
followed by viruses infecting bacteria (bacteriophages, Figure 2.4). This cycle results
FIGURE 2.4 Bacteriophage: A virus that infects bacteria.
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Bioprocessing of Viral Vaccines